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While some bandwidth could be recovered by putting a GUP on each subnet, the management of a large-scale GUP environment in a local LAN will likely take more time and effort than any nominal bandwidth savings.Recently we had a customer ask if all systems could be GUPs.Once clients have been installed and operating normally the definition updates are normally between 40kb-200kb.
In this scenario none of the agents will actually communicate with another GUP since a GUP can only retrieve updates from a SEPM.
While in the field we have seen clients use GUPs in different ways, the purpose of the GUPs was to reduce bandwidth requirements.
On a subnet over a WAN link, you would have a single client retrieving definitions from the SEPM.
Depending on how you publish definitions within your environment, something else to consider is the difference between cheap and expensive bandwidth.
In some environments client communication will go over the WAN while Internet traffic will traverse through a cheaper local ISP.